1. I Start With My Shutter Speed
For wildlife, shutter speed is one of the first things I think about because movement can change incredibly quickly.
Even when an animal appears still, it isn't necessarily still. An ear twitches. A bird turns its head. A bear shakes water from its fur. A wolf suddenly takes off running.
If I think there is any possibility of action, I would rather have more shutter speed than I need than discover afterward that I didn't have enough.
For general wildlife photography, I often want to be somewhere around 1/1000 second or faster. If I'm photographing birds in flight or fast action, I may be at 1/2000, 1/2500, or even faster.
But there isn't one magic shutter speed, and shutter speed doesn't exist in a vacuum. I'm also thinking about the available light and what that shutter speed is doing to my ISO.
If I'm photographing a bison standing quietly in beautiful early morning light, I may not need 1/2000 second. In that situation, I'm thinking about the slowest shutter speed I know is still fast enough for what the animal is doing. I don't want to unnecessarily push my ISO higher and give up image quality just for the sake of having a faster shutter speed.
At the same time, I don't want to push that shutter speed so low that I'm unprepared if the animal suddenly moves. It's a constant balance—enough shutter speed to make the photograph sharp, while keeping an eye on how hard I'm pushing the ISO in low light.
As the light improves, I have more room to increase my shutter speed. And if the animal's behavior changes, I'll make that adjustment immediately.
The important part is that I'm thinking about what the animal might do next—not just what it's doing at that exact moment.